林业公益性行业科研专项经费201504415;福建省自然科学基金项目2014J01137;福建农林大学科技创新专项基金CXZX2017141资助 This work was supported by the Forestry Scientific Research Projects in Public Interest of ChinaGrant 201504415;the Natural Science Foundation of Fujian Province,ChinaGrant 2014J01137;the Science and Technology Innovation Project of Fujian Agriculture and Forestry UniversityGrant CXZX2017141