林业公益性行业科研专项经费项目201504106;"十二五"农村领域国家科技计划项目2015BAD04B01资助 This work was supported by the Special Projects for Scientific Research in Forestry Public Welfare IndustryGrant 201504106;the National Science and Technology Planning Project for Rural Areas in the 12th Five-yearGrant 2015BAD04B01