资助项目国家重点基础研究发展计划项目2012CB955604,国家自然科学基金项目41275089、41305071、41425019、41661144016,海洋公益性行业科研专项201505013Funded by National Basic Research Program of ChinaGrant 2012CB955604;National Natural Science Foundation of ChinaGrants 51245089,41305071,41425019,and 41661144016;Public Science and Technology Research Funds Projects of OceanGrant 201505013