国家自然科学基金项目71473100;中组部青年拔尖人才支持计划项目;华中农业大学博士科研启动基金项目2662015BQ025.National Natural Science Foundation of China71473100;Youth Top-notch Talent Support Program of the Organization Department of the Central Committee of the CPC;Doctoral Scientific Research Foundation of Huazhong Agricultural University2662015BQ025