国家自然科学基金项目KAa17023A,31400552;福建农林大学杰出青年基金项目XJQ201613;福建农林大学国际科技合作与交流项目KXB16008AThe National Natural Science Foundation of ChinaKAa17023A, 31400552;Fujian Agriculture and Forestry University Funds for Distinguished Young ScholarXJQ201613;International Science and Technology Cooperation Program of Fujian Agriculture and Forestry UniversityKXB16008A