国家公益性行业农业科研专项201303143;江西省教育厅科学技术研究项目GJJ150435;国家现代肉牛牦牛产业技术体系项目CARS-38;国家星火项目2015GA730003.Project supported by the Special Scientific Research Fund of Agriculture Public Welfare Profession of ChinaGrant 201303143;Scientific Research Fund of Education Department of Jiangxi ProvinceGJJ150435;National Beef Cattle Industrial Technology System ProgramCARS-38;China Spark Program2015GA730003