国家自然科学基金项目31660672;现代农业产业技术体系资助项目CARS-38;江西省教育厅科学技术重点项目GJJ150376PProject supported by the Natural Science Foundation of China31660672;The modern agricultural industry technology systemCARS-38;Key project of science and technology of Jiangxi Provincial Education DepartmentGJJ150376