国家科技支撑计划项目2014BAK19B06-03;国家自然科学基金项目31270488,31501274;主要粮食作物产业化湖北省协同创新中心开放基金项目2015MS001 Project supported by National Science and tEchnology Support Program2014BAK19B06-03;the National Natural Science Foundation of China31270488,31501274;Hubei Collaborative Innovation Center for Grain Industry oPen Fund Project2015MS001