国家自然科学基金项目31000264、31470577;福建省高校产学合作科技重大项目2014N5005;福建农林大学林学院青年科学基金6112C039U Project supported by the National Natural Science Foundation of China31000264,31470577;Science and Technology Major Projects of Industry-University Cooperation in Universities and Colleges of Fujian Province2014N5005;Science Foundation for Youths in College of Forestry,Fujian Agriculture and Forestry Universi-ty6112C039U