湖北省自然科学基金项目2013CFB426;国家公益性行业农业科研专项201103016;国家油菜产业技术体系资助项目 CARS-13 Project supported by the Natural Science Foundation of Hubei Province,China2013CFB426;the Special Fund for Agro-scientific Research in the Public Interest201103016;the Industry Technology System of Rape-seed in ChinaCARS-13