中国水产科学研究院黄海水产研究所基本科研业务费20603022017001;国家自然科学基金41676148;泰山学者良种工程项目和现代农业产业技术体系专项资金CARS-48共同资助[This work was supported by Central Public-Interest Scientific Institution Basal Research Fund, Yellow Sea Fisheries Research Institute, CAFS20603022017001;National Natural Science Foundation of China41676148;Taishan Scholar Program for Seed Industry, and China Agriculture Research SystemCARS-48]