国家科技支撑计划课题2015BAD17B01;中国水产科学研究院黄海水产研究所基本科研业务费20603022016002共同资助[This work was supported by the National Science and Technology Support Plan Project2015BAD17B01;Special Scientific Research Funds for Central Non-Profit Institutes,Yellow Sea Fisheries Research Institute,Chinese Academy of Fishery Sciences20603022016002]