国家虾产业技术体系CARS-47;泰山产业领军人才项目LNJY2015002;青岛海洋科学与技术国家实验室鳌山科技创新计划项目2015ASKJ02共同资助[This work was supported by Modern Agro-industry Technology Research SystemCARS-47;the Program of Shandong Leading TalentLNJY2015002;Science and Technology Innovation Project of Qingdao National Laboratory for Marine Science and Technology 2015ASKJ02]