公益性行业农业科研专项201203017;青岛海洋科学与技术国家实验室鳌山科技创新计划项目2016ASKJ02;国家贝类产业技术体系CARS-47共同资助[This work was funded by Special Fund for Agro-Scientific Research in the Public Interest201203017;the Scientific and Technological Innovation Project financially supported by Qingdao National Laboratory for Marine Science and Technology2016ASKJ02;Modern Agro-Industry Technology Research System CARS-47]