国家鲆鲽类产业技术体系项目CARS-50;中国水产科学研究院黄海水产研究所级基本科研业务费20603022016003;国家国际科技合作专项项目2013DFA31410共同资助[This work was supported by China Agricultural Research SystemCARS-50;Special Scientific Research Funds for Central Non-Profit Institutes,Yellow Sea Fisheries Research Institute,Chinese Academy of Fishery Sciences20603022016003;International S&T Cooperation Program of China 2013DFA31410]