项目受国家自然科学基金.21675092,41476083,21275084,51525903;国家863计划2015AA034404;黄岛区海洋科技专项No.2014-4-1资助.Project supported by the National Natural Science Foundation of China.21675092,41476083,21275084,51525903;863 program2015AA034404;Marine Science and Technology Projects of Huangdao District2014-4-1