国家林业局引进国际先进林业科学技术项目No.2012-4-73;山东省自然科学基金No.ZR2013EEQ009;国家自然科学基金No.31270417 Supported by the State Forestry Administrational Foundation for the Introduction of International Advanced Forestry Technology No.2012-4-73,the Natural Science Foundation of Shandong ProvinceNo.ZR2013EEQ009 and the National Natural Science Foundation of ChinaNo.31270417