国家自然科学基金委-山东省人民政府联合资助海洋科学研究中心项目U1406404;国家重点基础研究发展计划973计划2015CB453301资助.Supported by the National Natural Science Foundation of China—Shandong Joint Fund for Marine Science Research Centers Under ContractU1406404;the National Program on Key Basic Research Project973 Program,2015CB453301