国家"十二五"农村领域科技计划项目2012AA101204-2;海南省高层次创新创业人才启动基金;中央级公益性科研院所基本科研业务费专项1630032013005 [Supported by National Scientific and Technological Programs in Rural Fields2012AA101204-2;the Initial Fund of High-levele Creative Talents in Hainan Province;the National Nonprofit Institute Research Grant of CATAS-TCGRI1630032013005]