国家自然科学基金31160137;广西自然科学基金2015GXNSFBA139051, 2016GXNSFAA380122;广西植物研究所基本业务费项目桂植业 14004 [ Supported by the National Natural Science Foundation of China31160137;the Natural Science Foundation of Guangxi2015GXNSFBA139051, 2016GXNSFAA380122;the Science Research Foundation of Guangxi Institute of Botany Guizhiye 14004]