国家自然科学基金31201191,31660386;江西省自然科学基金20132BAB214009,20142BAB204012;江西省教育厅项目GJJ14248;江西省亚热带植物资源保护与利用重点实验室开放基金YRD201403[Supported by the National Natural Science Foundation of China 31201191,31660386;Natural Science Foundation of Jiangxi 20132BAB214009,20142BAB204012; Jiangxi Program of Education Office GJJ14248;Open Fund of Jiangxi Key Laboratory of Protection and Utilization of Subtropical Plant Resources YRD201403]。