美国国家科学基金INT-0204923、EAR-0716235;中国国家自然科学基金重点项目41430102;中国科学院战略性先导科技专项编号:XDB03020104资助.This research was funded by the U.S.National Science FoundationINT-0204923 and EAR-0716235 to Wang;the National Natural Science Foundation of China41430102 to Deng;the Chinese Academy of Sciences XDB03020104 to Deng.We are grateful to Dr.Penny Higgins and Dr.Bruce MacFadden for helpful comments and suggestions that greatly improved this manuscript