国家自然科学基金40872019;the grants from the National Natural Science Foundation of China40872019;from the Major Basic Research Projects2006CB806400;the Ministry of Science and Technology of China to SQH,WXC and LC;from the Canadian Museum of NatureR34;from the National Natural Science Foundation40772015;40121202;the CAS/SAFE A International Partnership Program for Creative Research Teams to LC and WXC