全军后勤科研“十二五”计划面上项目CWS12J017;国家自然科学基金面上项目81271875;美国NIH重大国际合作课题U19A1089672;Supported by the General Program of Military Scientific Research of China CWS12J017,the General Program of National Natural Science Foundation of China 81271875 and the Major Internation Coorperation Project of National Institutes of Health,USA U19AI089672.