公益性行业农业科研专项201203065[This study was supported by the Special Fund for Agro-Scientific Research in the Public Interest 201203065];国家自然科学基金项目31472280[the National Natural Science Foundation of China 31472280];高等学校博士学科点专项科研基金20123104110006 the Ministry of Education's Doctoral Discipline Foundation 20123104110006