国家自然科学基金项目31300538,31400540the National Natural Science Foundation of China Grant No.31300538 and 31400540;西北农林科技大学基本科研业务费专项资金编号:QN2013082Basic Scientific Research Business Expenses of Northwest A&F UniversityGrant No.QN2013082;西北农林科技大学第二批基本科研业务费青年培育专项2452015335Youth Development Project of Second Basic Scientific Research Business Expenses of Northwest A&F University Grant No.2452015335